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Living underground in Giha Village, Simon and Kamina dream of a life on the surface. When their home is invaded from above, it is up to them to activate the mysterious mecha Lagann and save their village.

Living underground in Giha Village, Simon and Kamina dream of a life on the surface. When their home is invaded from above, it is up to them to activate the mysterious mecha Lagann and save their village.

Now that they've made it to the surface, Simon and Kamina must join together with Yoko and her village to defeat the Beastmen pursuing them.

Now that they've made it to the surface, Simon and Kamina must join together with Yoko and her village to defeat the Beastmen pursuing them.

Kamina and Simon must work together to defeat a dangerous new Beastman threat.

Kamina and Simon must work together to defeat a dangerous new Beastman threat.

Team Gurren must work together with the eccentric Black Syblings to take down a Beastman foe on empty stomachs.

Team Gurren must work together with the eccentric Black Syblings to take down a Beastman foe on empty stomachs.

Team Gurren must find their way out of a strange underground village where they worship Gunmen as divine beings.

Team Gurren must find their way out of a strange underground village where they worship Gunmen as divine beings.

Continuing to the Gunmen rallying point, Team Gurren finds themselves on a strange hot springs getaway.

Continuing to the Gunmen rallying point, Team Gurren finds themselves on a strange hot springs getaway.

Thymilph, one of the Four Divine Generals of the mysterious Spiral King, arrives to support Viral's Enki, with the massive mobile Gunmen fortress, the Dai-Ganzan. Kamina and Simon pilot the Gurren-Lagann against seemingly overwhelming odds.

Thymilph, one of the Four Divine Generals of the mysterious Spiral King, arrives to support Viral's Enki, with the massive mobile Gunmen fortress, the Dai-Ganzan. Kamina and Simon pilot the Gurren-Lagann against seemingly overwhelming odds.

Having temporarily stalled Viral and Thymilph in the last skirmish, the Team Gurren and their newly arrived allies join forces as "Team Dai-Gurren" and launch a decisive strike against the regrouping Gunmen force.

Having temporarily stalled Viral and Thymilph in the last skirmish, the Team Gurren and their newly arrived allies join forces as "Team Dai-Gurren" and launch a decisive strike against the regrouping Gunmen force.

Team Dai-Gurren is still mourning on Kamina's death. Simon notices a Gunman throw a capsule into a deep valley. He inspects it and is able to open it, and inside is a human girl named Nia, but their introduction is cut short when an enemy Gunman appears.

Team Dai-Gurren is still mourning on Kamina's death. Simon notices a Gunman throw a capsule into a deep valley. He inspects it and is able to open it, and inside is a human girl named Nia, but their introduction is cut short when an enemy Gunman appears.

Having been temporarily spared by Adiane, another of the Four Divine Generals, Team Dai-Gurren uses the reprieve to reassess their situation. Nia learns of Kamina's significance in Simon's erratic behavior and offers him some stinging, yet honest advice.

Having been temporarily spared by Adiane, another of the Four Divine Generals, Team Dai-Gurren uses the reprieve to reassess their situation. Nia learns of Kamina's significance in Simon's erratic behavior and offers him some stinging, yet honest advice.

Team Dai-Gurren falls into a trap set by the third member of the Four Divine Generals, Guame. The Team is taken captive and stored underground to await execution. While digging an escape path, Simon finally understands Kamina's last words to him.

Team Dai-Gurren falls into a trap set by the third member of the Four Divine Generals, Guame. The Team is taken captive and stored underground to await execution. While digging an escape path, Simon finally understands Kamina's last words to him.

On their way to Teppelin, the Beastmen's capital city, Team Dai-Gurren stumbles upon a vast ocean and must wait for modifications on the Dai-Gurren to be completed before crossing it. While the rest of the group play at the beach, Yoko feels uneasy.

On their way to Teppelin, the Beastmen's capital city, Team Dai-Gurren stumbles upon a vast ocean and must wait for modifications on the Dai-Gurren to be completed before crossing it. While the rest of the group play at the beach, Yoko feels uneasy.

Trying to do her part in Team Dai-Gurren, Nia decides to become a cook, but just before the crew have a taste of her dishes, Cytomander, the fourth member of the Divine Generals, launches an aerial attack on the Dai-Gurren. Simon leads the counter-attack.

Trying to do her part in Team Dai-Gurren, Nia decides to become a cook, but just before the crew have a taste of her dishes, Cytomander, the fourth member of the Divine Generals, launches an aerial attack on the Dai-Gurren. Simon leads the counter-attack.

At last, Team Dai-Gurren arrives at the outskirts of Teppelin, but before aiming to the city, they must face a combined full-scale attack of Cytomander and Guame's forces.

At last, Team Dai-Gurren arrives at the outskirts of Teppelin, but before aiming to the city, they must face a combined full-scale attack of Cytomander and Guame's forces.

The last of the Divine Generals fall, and the city of Teppelin collapses, revealing a Gunman of gigantic proportions. Simon and Nia struggle to pass through it in order to confront her father, the Spiral King himself.

The last of the Divine Generals fall, and the city of Teppelin collapses, revealing a Gunman of gigantic proportions. Simon and Nia struggle to pass through it in order to confront her father, the Spiral King himself.

A recap of the story's events, as of episode 15.

A recap of the story's events, as of episode 15.

During the following seven years, a new civilization rises and prospers under Simon's leadership. However, these peaceful times come to end when the threat mentioned in Lordgenome's last words starts to reveal itself.

During the following seven years, a new civilization rises and prospers under Simon's leadership. However, these peaceful times come to end when the threat mentioned in Lordgenome's last words starts to reveal itself.

After the Anti-Spirals' first attack and Nia's astonishing announcement, Simon looks desperately for her, while Rossiu finally completes his secret project.

After the Anti-Spirals' first attack and Nia's astonishing announcement, Simon looks desperately for her, while Rossiu finally completes his secret project.

To soothe the public opinion, Rossiu overthrows Simon and puts him under arrest. But when the Anti-Spirals launch a large-scale attack on Kamina City, he is left with no option but to let Simon engage the enemy using the Gurren-Lagann.

To soothe the public opinion, Rossiu overthrows Simon and puts him under arrest. But when the Anti-Spirals launch a large-scale attack on Kamina City, he is left with no option but to let Simon engage the enemy using the Gurren-Lagann.

As Simon arrives in prison, he unexpectedly meets Viral, who comments on how far he has fallen. Meanwhile, the truth about Rossiu's emergency plan and the consequences for those left behind on Earth are revealed.

As Simon arrives in prison, he unexpectedly meets Viral, who comments on how far he has fallen. Meanwhile, the truth about Rossiu's emergency plan and the consequences for those left behind on Earth are revealed.

One year earlier, Yoko starts working as a schoolteacher on an island until she learns of the situation at Teppelin and returns to break Simon free. Once again reunited, Simon and the members of Team Dai-Gurren launch themselves into space.

One year earlier, Yoko starts working as a schoolteacher on an island until she learns of the situation at Teppelin and returns to break Simon free. Once again reunited, Simon and the members of Team Dai-Gurren launch themselves into space.

The Gurren-Lagann and the Arc-Gurren join together to form the massive Arc-Gurren-Lagann and stop the Human Annihilation System.

The Gurren-Lagann and the Arc-Gurren join together to form the massive Arc-Gurren-Lagann and stop the Human Annihilation System.

The members of Team Dai-Gurren make their preparations for the imminent war against the Anti-Spirals, but Rossiu is missing. However, when Kinon tells Simon about what he really intends to do, both start to look desperately for him.

The members of Team Dai-Gurren make their preparations for the imminent war against the Anti-Spirals, but Rossiu is missing. However, when Kinon tells Simon about what he really intends to do, both start to look desperately for him.

Guided by Simon and Nia's feelings for each other, Team Dai-Gurren break through dimensions in search for the Anti-Spirals' homeworld. But after falling into the enemy's trap, they end up losing some of their members, due to its vicious offensive.

Guided by Simon and Nia's feelings for each other, Team Dai-Gurren break through dimensions in search for the Anti-Spirals' homeworld. But after falling into the enemy's trap, they end up losing some of their members, due to its vicious offensive.

Team Dai-Gurren struggles to escape from the ocean of despair created by the Anti-Spirals without success, until history repeats itself, as another irreplaceable member of the Team sacrifices himself to save the others.

Team Dai-Gurren struggles to escape from the ocean of despair created by the Anti-Spirals without success, until history repeats itself, as another irreplaceable member of the Team sacrifices himself to save the others.

After finally transforming their flagship into the Super Galactic Gurren-Lagann, Simon and his friends successfully fight back against the enemy. When their enemy starts to use their own memories and desires against them, Kamina reappears to help.

After finally transforming their flagship into the Super Galactic Gurren-Lagann, Simon and his friends successfully fight back against the enemy. When their enemy starts to use their own memories and desires against them, Kamina reappears to help.

At last, Simon and Nia are reunited, and by combining the power of its members, Team Dai-Gurren creates the galaxy-sized Tengen Toppa Gurren-Lagann, and engage the Anti-Spiral threat in a decisive battle to determine the fate of the universe.

At last, Simon and Nia are reunited, and by combining the power of its members, Team Dai-Gurren creates the galaxy-sized Tengen Toppa Gurren-Lagann, and engage the Anti-Spiral threat in a decisive battle to determine the fate of the universe.